生活型

herb

形態特徵

• Perennial herbs with stolons; stems erect, ascending or prostrate, puberulent at first, glabrescent toward base.
• Leaves equally to unequally paired, membranaceous to chartaceous, somewhat puberulent or with conical trichomes to nearly glabrous above and along nerves beneath, rhombic-ovate, ovate to ovate-lanceolate, or narrowly elliptic to elliptic, 1.5-17 cm long, 1-6.5 cm wide, apex obtuse to acuminate, with an obscurely serrulate acumine, margins serrate to crenate-serrate, base broadly cuneate to obtuse, 3-nerved; petioles 0.6-6.0 cm long, puberulent or with conical trichomes; stipules 2, connate at base, cordiform, to 5 mm long, 6.4 mm wide.
• Male inflorescences axillary, cymosely to paniculately arranged glomerules; peduncles to 5.1 cm long. Female inflorescences axillary, short cymosely arranged glomerules; peduncles to 1 cm long. Male flowers usually brownish red or deep red on upper parts, laterally compressed, obovoid; sepals 4, subequal, connate halfway, cymbiform; stamens 4; pistil rudimentary. Female flowers oblong-ovoid; sepals 3, unequal; staminodes 3; ovary oblong-ovoid.
• Achenes tuberculate, laterally compressed, obliquely ellipsoid.

物候

Flowering and fruiting December to June, or to July at higher altitudes.

分布

• 台灣 : throughout between 400-2,600 m, small ravines and moist forest edges and forest floors.
• 世界 : S. mainland China, Ryukyus and Vietnam.

引證標本

TAIPEI: Yang-Chin Highway, Shih 3412. ILAN: Nanshantsun, Shih 2869; Su-Hua Highway, Shih 2840. TAOYUAN: Rarashan, Chen s. n. 1994. HSINCHU: Wuchihshan, Ito s. n. 1914. MIAOLI: Nanchuang, Shimada s. n. 1923. TAICHUNG: Pilushan Logging Trail, Shih 3022. NANTOU: Chitou, Ou & Kao 8909; Meifong, Shih 2942; Tsueifong, Shih 2932. CHIAYI: Alishan-Erwanping, Huang 5026; Funkiko to Toloma, Hayata s. n. 1914. TAITUNG: Chihpen Logging Trail, Shih 2650; Lotachienshan, Shih 2580. HUALIEN: Tienhsiang, Shimizu & Kao 10613.

引用文獻

• YANG, YUEN-PO, SHIH, BING-LING and LIU, HO-YIH, Flora of Taiwan, second edition 2: 236. 1996.  (Fl. Taiwan 2nd edit., 台灣植物誌第二版)